3 分类类型
属
5 甲型杆状病毒属成员
| 种 |
| 双线盗毒蛾核型多角体病毒(Porthesia scintillans nucleopolyhedrovirus) |
| 落叶松叶蜂核型多角体病毒(Pristiphora erichsonii nucleopolyhedrovirus) |
| 芩叶蜂核型多角体病毒(Pristiphora geniculata nucleopolyhedrovirus) |
| 斜纹夜蛾核型多角体病毒(Prodenia litosia nucleopolyhedrovirus) |
| 黄纹夜蛾核型多角体病毒(Prodenia praefica nucleopolyhedrovirus) |
| 高原斜纹夜蛾核型多角体病毒(Prodenia terricola nucleopolyhedrovirus) |
| 虚线尺蛾核型多角体病毒(Protoboarmia porcelaria nucleopolyhedrovirus) |
| 变异黏虫核型多角体病毒(Pseudaletia convecta nucleopolyhedrovirus) |
| 粘虫核型多角体病毒(Pseudaletia separata nucleopolyhedrovirus) |
| 大豆尺蠖蛾核型多角体病毒(Pseudoplusia includens nucleopolyhedrovirus) |
| 异角牧草蛾核型多角体病毒( Pterolocera amplicornisnucleopolyhedrovirus) |
| 落叶松卷蛾核型多角体病毒(Ptycholomoides aeriferana nucleopolyhedrovirus) |
| 顺脉小尺蠖核型多角体病毒(Ptychopoda seriata nucleopolyhedrovirus) |
| 杨扇舟蛾核型多角体病毒(Pygaera anastomosis nucleopolyhedrovirus) |
| 分月扇舟蛾核型多角体病毒(Pygaera fulgurita nucleopolyhedrovirus) |
| 野螟核型多角体病毒(Pyrausta diniasalis nucleopolyhedrovirus) |
| 薄荷灰夜蛾核型多角体病毒(Rachiplusia ou nucleopolyhedrovirus) |
| 夏梢小卷蛾核型多角体病毒 (Rhyacionia duplana nucleopolyhedrovirus) |
| 安哥拉黑翅菌蝇核型多角体病毒( Rhynchosciara angelae nucleopolyhedrovirus) |
| 铜色翅菌蝇核型多角体病毒(Rhynchosciara hollaenderi nucleopolyhedrovirus) |
| 米勒黑翅菌蝇核型多角体病毒(Rhynchosciara milleri nucleopolyhedrovirus) |
| 桑蟥蚕蛾核型多角体病毒(Rondotia menciana nucleopolyhedrovirus) |
| 樗树大蚕核型多角体病毒(Samia cynthia nucleopolyhedrovirus) |
| 樗天蚕核型多角体病毒(Samia pryeri nucleopolyhedrovirus) |
| 蓖麻蚕核型多角体病毒(Samia ricini nucleopolyhedrovirus) |
| 褐带卷蛾核型多角体病毒 (Adoxophyes honmai NPV) |
| 小地老虎核型多角体病毒(Agrotis ipsilon MNPV) |
| 黎豆夜蛾多衣壳核型多角体病毒 (Anticarsia gemmatalis MNPV) |
| 苜蓿银纹夜蛾核型多角体病毒(Autographa californica MNPV) |
| 家蚕核型多角体病毒(Bombyx mori NPV) |
| 油桐尺蠖核型多角体病毒 (Buzura suppressaria NPV) |
| 云杉卷叶蛾核型多角体病毒DEF型(Choristoneura fumiferana DEF MNPV) |
| 云杉卷叶蛾核型多角体病毒(Choristoneura fumiferana NPV) |
| 蔷薇卷叶蛾核型多角体病毒(Choristoneura rosaceana MNPV) |
| 茶尺蠖核型多角体病毒(Ecotropis oblique NPV) |
| 苹果浅棕色夜蛾核型多角体病毒(Epiphyas postivittana NPV) |
| 棉铃虫核型多角体病毒(Helicoverpa armigera NPV) |
| 美洲棉铃虫核型多角体病毒(Helicoverpa zea SNPV) |
| 舞毒蛾核型多角体病毒(Lymantria dispar MNPV) |
| 甘蓝夜蛾核型多角体病毒(Mamestra brassicae MNPV) |
| 蓓带夜蛾核型多角体病毒A型(Mamestra configurata NPV-A) |
| 蓓带夜蛾核型多角体病毒B型(Mamestra configurata NPV-B) |
| 黄杉毒蛾核型多角体病毒(Orgyia pseudotsugata MNPV) |
| 甜菜夜蛾核型多角体病毒(Spodoptera exigua MNPV) |
| 草地贪夜蛾核型多角体病毒(Spodoptera frugiperda MNPV) |
| 海灰翅夜蛾核型多角体病毒(Spodoptera littoralis NPV) |
| 斜纹夜蛾核型多角体病毒(Spodoptera litura NPV) |
| 埃塞克斯弄蝶核型多角体病毒 (Thymelicus lineola NPV) |
| 粉纹夜蛾单核衣壳核型多角体病毒(Trichoplusia ni SNPV) |
| 蝙蝠蛾核型多角体病毒(Wiseana signata NPV) |
| 美国白蛾核型多角体病毒 (Hyphantria cunea NPV) |
| 黄杉毒蛾单核衣壳核型多角体病毒(Orgyia pseudotsugata SNPV) |
| 草虾核型多角体病毒(Penaeus monodon NPV) |
| 松美蛾核型多角体病毒 (Panolis flammea NPV) |
| 醋栗尺蠖核型多角体病毒(Abraxas grossulariata nucleopolyhedrovirus) |
| 蓖麻夜蛾核型多角体病毒(Achaeas janata nucleopolyhedrovirus) |
| 小蜡螟核型多角体病毒(Achroia grisella nucleopolyhedrovirus) |
| 韧皮尺蠖核型多角体病毒(Acidalia carticcaria nucleopolyhedrovirus) |
| 西部黑头长翅卷蛾核型多角体病毒(Acleris gloverana nucleopolyhedrovirus) |
| 东部黑头长翅卷叶蛾核型多角体病毒(Acleris variana nucleopolyhedrovirus) |
| 锐剑纹夜蛾核型多角体病毒(Acronycta aceris nucleopolyhedrovirus) |
| 黑行军虫核型多角体病毒(Actebia fennica nucleopolyhedrovirus) |
| 柳天蚕核型多角体病毒(Actias selene nucleopolyhedrovirus) |
| 曙夜蛾核型多角体病毒(Adisura atkinsoni nucleopolyhedrovirus) |
| 茶小卷叶蛾核型多角体病毒(Adoxophyes orana nucleopolyhedrovirus) |
| 甘薯烦夜蛾核型多角体病毒(Aedia leucomelas nucleopolyhedrovirus) |
| 龟甲蝶核型多角体病毒(Aglais urticae nucleopolyhedrovirus) |
| 香子兰蝶核型多角体病毒(Agraulis vanillae nucleopolyhedrovirus) |
| 警纹地老虎核型多角体病毒(Agrotis exclamationis nucleopolyhedrovirus) |
| 小地老虎核型多角体病毒(Agrotis ipsilon nucleopolyhedrovirus) |
| 黄地老虎核型多角体病毒(Agrotis segetum nucleopolyhedrovirus) |
| 棉叶波纹夜蛾核型多角体病毒(Alabama argillacea nucleopolyhedrovirus) |
| 小壁板夜蛾核型多角体病毒(Aletia oxygala nucleopolyhedrovirus) |
| 粉白灯蛾核型多角体病毒(Alphaea phasma nucleopolyhedrovirus) |
| 秋星尺蠖核型多角体病毒(Alsophila pometaria nucleopolyhedrovirus) |
| 八字地老虎核型多角体病毒(Amathes c-nigrum nucleopolyhedrovirus) |
| 葡萄天蛾核型多角体病毒(Amphelophaga rubiginosa nucleopolyhedrovirus) |
| 裂尺蛾核型多角体病毒(Amphidasis cognataria nucleopolyhedrovirus) |
| 花生灯蛾核型多角体病毒(Amsacta albistriga nucleopolyhedrovirus) |
| 红缘灯蛾核型多角体病毒(Amsacta lactinea nucleopolyhedrovirus) |
| 桑灯蛾核型多角体病毒(Amsacta moorei nucleopolyhedrovirus) |
| 柑橘卷叶蛾核型多角体病毒(Amyelois transitella nucleopolyhedrovirus) |
| 葫芦夜蛾核型多角体病毒(Anadevidia peponis nucleopolyhedrovirus) |
| 地中海粉螟核型多角体病毒(Anagasta kuehniella nucleopolyhedrovirus) |
| 尺蠖核型多角体病毒(Anaitis plaguata nucleopolyhedrovirus) |
| 栎橙纹犀额蛾核型多角体病毒(Anisota senatoria nucleopolyhedrovirus) |
| 棉小造桥虫核型多角体病毒(Anomis flava nucleopolyhedrovirus) |
| 印黄檀造桥虫核型多角体病毒(Anomis sabulifera nucleopolyhedrovirus) |
| 八字夜蛾核型多角体病毒(Anomogyna elimata nucleopolyhedrovirus) |
| 软羽澳蛾核型多角体病毒(Anthela varia nucleopolyhedrovirus) |
| 蛀澳蛾核型多角体病毒(Anthelia hyperborea nucleopolyhedrovirus) |
| 塔萨大蚕蛾核型多角体病毒(Antheraea paphia nucleopolyhedrovirus) |
| 柞蚕核型多角体病毒(Antheraea pernyi nucleopolyhedrovirus) |
| 北美天蚕蛾核型多角体病毒(Antheraea polyphemus nucleopolyhedrovirus) |
| 日本柞蚕核型多角体病毒(Antheraea yamamai nucleopolyhedrovirus) |
| 墨西哥棉铃象核型多角体病毒(Anthonomus grandis nucleopolyhedrovirus) |
| 标本皮蠹核型多角体病毒(Anthrenus museorum nucleopolyhedrovirus) |
| 褐污秀夜蛾核型多角体病毒(Apamea anceps nucleopolyhedrovirus) |
| 沙枣尺蠖核型多角体病毒(Apocheima cinerarius nucleopolyhedrovirus) |
| 毛茸枣尺蠖 核型多角体病毒(Apocheima pilosaria nucleopolyhedrovirus) |
| 山楂粉蝶核型多角体病毒(Aporia crataegi nucleopolyhedrovirus) |
| 钩麦蛾核型多角体病毒(Aproaerema modicella nucleopolyhedrovirus) |
| 花斑蛱蝶核型多角体病毒(Araschnia levana nucleopolyhedrovirus) |
| 巢黄卷蛾核型多角体病毒(Archips cerasivoranus nucleopolyhedrovirus) |
| 豹灯蛾核型多角体病毒(Arctia caja nucleopolyhedrovirus) |
| 绒毛灯蛾核型多角体病毒(Arctia villica nucleopolyhedrovirus) |
| 灯蛾核型多角体病毒(Ardices glatignyi nucleopolyhedrovirus) |
| 桦三节叶蜂核型多角体病毒(Arge pectoralis nucleopolyhedrovirus) |
| 豹蛱蝶核型多角体病毒(Argynnis paphia nucleopolyhedrovirus) |
| 基纹银纹夜蛾核型多角体病毒(Argyrogramma basigera nucleopolyhedrovirus) |
| 阿斯特勒蝶核型多角体病毒(Astero campaceltis nucleopolyhedrovirus) |
| 生菜尺蠖核型多角体病毒(Autographa biloha nucleopolyhedrovirus) |
| 双斑夜蛾核型多角体病毒 (Autographa bimaculata nucleopolyhedrovirus) |
| 汉马夜蛾核型多角体病毒(Autographa gamma nucleopolyhedrovirus) |
| 黑点银纹夜蛾核型多角体病毒 (Autographa nigrisigna nucleopolyhedrovirus) |
| 分脉夜蛾核型多角体病毒(Autographa precationis nucleopolyhedrovirus) |
| 云斑天牛核型多角体病毒(Batocera lineolata nucleopolyhedrovirus) |
| 白尾螟蛾核型多角体病毒(Bellura gortynoides nucleopolyhedrovirus) |
| 柳枯叶蛾核型多角体病毒(Bhima undulosa nucleopolyhedrovirus) |
| 桦尺蠖核型多角体病毒(Biston betularia nucleopolyhedrovirus) |
| 多毛尺蠖核型多角体病毒(Biston hirtaria nucleopolyhedrovirus) |
| 黄角尺蠖核型多角体病毒(Biston hispidaria nucleopolyhedrovirus) |
| 油茶尺蠖核型多角体病毒(Biston marginata nucleopolyhedrovirus) |
| 褐纹大尺蠖核型多角体病毒(Biston robustum nucleopolyhedrovirus) |
| 杨尺蠖核型多角体病毒(Biston strataria nucleopolyhedrovirus) |
| 双尖顶尺蠖核型多角体病毒(Boarmia bistortata nucleopolyhedrovirus) |
| 茶尺蠖核型多角体病毒(Boarmia obliqua nucleopolyhedrovirus) |
| 棉潜蛾核型多角体病毒(Bucculatrix thurbeliella nucleopolyhedrovirus) |
| 松天蛾核型多角体病毒(Bupalus piniarius nucleopolyhedrovirus) |
| 云尺蠖核型多角体病毒(Buzura thibtaria nucleopolyhedrovirus) |
| 粉斑螟核型多角体病毒(Cadra cautella nucleopolyhedrovirus) |
| 葡萄干果斑螟核型多角体病毒(Cadra figulilella nucleopolyhedrovirus) |
| 黑颊丽蝇核型多角体病毒(Calliphora vomitoria nucleopolyhedrovirus) |
| 标冬夜蛾核型多角体病毒(Calophasia lunula nucleopolyhedrovirus) |
| 桑蓑蛾核型多角体病毒(Canephora asiatica nucleopolyhedrovirus) |
| 云杉灰尺蠖核型多角体病毒(Caripeta divisata nucleopolyhedrovirus) |
| 桃小食心虫核型多角体病毒(Carposina niponensis nucleopolyhedrovirus) |
| 乳夜蛾核型多角体病毒(Catabena esula nucleopolyhedrovirus) |
| 连裳夜蛾核型多角体病毒(Catocala conjuncta nucleopolyhedrovirus) |
| 睡莲裳夜蛾核型多角体病毒(Catocala nymphaea nucleopolyhedrovirus) |
| 阿戈戈裳夜蛾核型多角体病毒(Catocala nymphagoga nucleopolyhedrovirus) |
| 牛角果粉蝶核型多角体病毒(Catopsilia Pomona nucleopolyhedrovirus) |
| 云杉扁叶蜂核型多角体病毒(Cephalcia abietis nucleopolyhedrovirus) |
| 斑马纹夜蛾核型多角体病毒(Ceramica picta nucleopolyhedrovirus) |
| 豆叶行军虫核型多角体病毒(Ceramica pisi nucleopolyhedrovirus) |
| 牧草夜蛾核型多角体病毒(Cerapteryx graminis nucleopolyhedrovirus) |
| 杨二尾舟蛾核型多角体病毒(Cerura hermelina nucleopolyhedrovirus) |
| 二化螟核型多角体病毒(Chilo suppressalis nucleopolyhedrovirus) |
| 柳色卷蛾核型多角体病毒(Choristoneura conflicatana nucleopolyhedrovirus) |
| 异色卷蛾核型多角体病毒(Choristoneura diversana nucleopolyhedrovirus) |
| 紫色卷蛾核型多角体病毒(Choristoneura murinana nucleopolyhedrovirus) |
| 西枞色卷蛾核型多角体病毒(Choristoneura occidentalis nucleopolyhedrovirus) |
| 松色卷蛾核型多角体病毒(Choristoneura pinus nucleopolyhedrovirus) |
| 玫瑰色卷蛾核型多角体病毒(Choristoneura rosaceana nucleopolyhedrovirus) |
| 锞纹夜蛾核型多角体病毒(Chrysodeixis chalcites nucleopolyhedrovirus) |
| 南方锞纹夜蛾核型多角体病毒(Chrysodeixis eriosoma nucleopolyhedrovirus) |
| 草蛉核型多角体病毒(Chrysopa perla nucleopolyhedrovirus) |
| 链斑尺蠖核型多角体病毒(Cingilia caternaria nucleopolyhedrovirus) |
| 黄刺蛾核型多角体病毒(Cnidocampa flavescens nucleopolyhedrovirus) |
| 落叶松鞘蛾核型多角体病毒(Coleophora laricella nucleopolyhedrovirus) |
| 橙色豆粉蝶核型多角体病毒(Colias electo nucleopolyhedrovirus) |
| 苜蓿粉蝶核型多角体病毒(Colias eurytheme nucleopolyhedrovirus) |
| 雷斯波粉蝶核型多角体病毒(Colias lesbia nucleopolyhedrovirus) |
| 云纹粉蝶核型多角体病毒(Colias philodice nucleopolyhedrovirus) |
| 粉花凌霄大蚕蛾核型多角体病毒(Coloradia pandora nucleopolyhedrovirus) |
| 米蛾核型多角体病毒(Corcyra cephalonica nucleopolyhedrovirus) |
| 芦苇枯叶蛾核型多角体病毒(Cosmotriche potatoria nucleopolyhedrovirus) |
| 芳香木蠹蛾核型多角体病毒(Cossus cossus nucleopolyhedrovirus) |
| 落叶松隐斑螟核型多角体病毒(Cryptoblabes lariciana nucleopolyhedrovirus) |
| 非金合欢袋蛾核型多角体病毒(Cryptothelea junodi nucleopolyhedrovirus) |
| 大袋蛾核型多角体病毒(Cryptothelea variegata nucleopolyhedrovirus) |
| 木撩尺蠖核型多角体病毒(Culcula panterinaria nucleopolyhedrovirus) |
| 波纹杂毛虫核型多角体病毒(Cyclophragma undans nucleopolyhedrovirus) |
| 双斑杂毛虫核型多角体病毒(Cyclophrama yamadai nucleopolyhedrovirus) |
| 苹果小卷蛾核型多角体病毒(Cydia pomonella nucleopolyhedrovirus) |
| 杉茸毒蛾核型多角体病毒(Dasychira abietis nucleopolyhedrovirus) |
| 柳杉茸毒蛾核型多角体病毒(Dasychira argentata nucleopolyhedrovirus) |
| 松茸毒蛾核型多角体病毒(Dasychira axutha nucleopolyhedrovirus) |
| 黄茸毒蛾核型多角体病毒(Dasychira basiflava nucleopolyhedrovirus) |
| 相思茸毒蛾核型多角体病毒(Dasychira confusa nucleopolyhedrovirus) |
| 蔚茸毒蛾核型多角体病毒(Dasychira glaucinoptera nucleopolyhedrovirus) |
| 富毒蛾核型多角体病毒(Dasychira locuples nucleopolyhedrovirus) |
| 沁茸毒蛾核型多角体病毒(Dasychira mendosa nucleopolyhedrovirus) |
| 云杉茸毒蛾核型多角体病毒(Dasychira plagiata nucleopolyhedrovirus) |
| 拟杉茸毒蛾核型多角体病毒(Dasychira pseudabietis nucleopolyhedrovirus) |
| 茸毒蛾核型多角体病毒(Dasychira pudibunda nucleopolyhedrovirus) |
| 白腰天蛾核型多角体病毒(Deilephiila elpenor nucleopolyhedrovirus) |
| 枞灰尺蠖核型多角体病毒(Deileptenia ribeata nucleopolyhedrovirus) |
| 云南松毛虫核型多角体病毒(Dendrolimus latipennis nucleopolyhedrovirus) |
| 欧洲松毛虫核型多角体病毒(Dendrolimus pini nucleopolyhedrovirus) |
| 马尾松毛虫核型多角体病毒(Dendrolimus punctatus nucleopolyhedrovirus) |
| 赤松毛虫核型多角体病毒(Dendrolimus spectabilis nucleopolyhedrovirus) |
| 火腿皮蠹核型多角体病毒(Dermestes lardarius nucleopolyhedrovirus) |
| 金杂翅夜蛾核型多角体病毒(Diachrysia orichalcea nucleopolyhedrovirus) |
| 尘污灯蛾核型多角体病毒(Diacrisia obliqua nucleopolyhedrovirus) |
| 紫红污灯蛾核型多角体病毒(Diacrisia purpurata nucleopolyhedrovirus) |
| 黄毛污灯蛾核型多角体病毒(Diacrisia virginica nucleopolyhedrovirus) |
| 小玻灯蛾核型多角体病毒(Diaphora mendica nucleopolyhedrovirus) |
| 巨座玉米螟核型多角体病毒(Diatraea grandiosella nucleopolyhedrovirus) |
| 小蔗螟核型多角体病毒(Diatraea saccharalis nucleopolyhedrovirus) |
| 桃蛀螟核型多角体病毒(Dichocrosis punctiferalis nucleopolyhedrovirus) |
| 银杏大蚕蛾核型多角体病毒(Dictyoploca japonica nucleopolyhedrovirus) |
| 巢大蚕蛾核型多角体病毒(Dicycla oo nucleopolyhedrovirus) |
| 无翅虫核型多角体病毒(Dilta hibernica nucleopolyhedrovirus) |
| 伪节梢斑螟核型多角体病毒(Dioryctria pseudotsugella nucleopolyhedrovirus) |
| 赤棉铃虫核型多角体病毒(Diparopsis watersi nucleopolyhedrovirus) |
| 欧洲云杉锯角叶蜂核型多角体病毒(Diprion hercyniae nucleopolyhedrovirus) |
| 六万松锯角叶蜂核型多角体病毒(Diprion liuwanensis nucleopolyhedrovirus) |
| 松黑点锯角叶蜂核型多角体病毒(Diprion nipponica nucleopolyhedrovirus) |
| 苍白色锯角叶蜂核型多角体病毒(Diprion pallida nucleopolyhedrovirus) |
| 红排锯角叶蜂核型多角体病毒(Diprion pindrowi nucleopolyhedrovirus) |
| 松大锯角叶蜂核型多角体病毒(Diprion pini nucleopolyhedrovirus) |
| 云杉锯角叶蜂核型多角体病毒(Diprion polytoma nucleopolyhedrovirus) |
| 五针松锯角叶蜂核型多角体病毒(Diprion similis nucleopolyhedrovirus) |
| 蚕蛾核型多角体病毒(Dirphia gragatus nucleopolyhedrovirus) |
| 栗刺蛾核型多角体病毒(Doratifera casta nucleopolyhedrovirus) |
| 黑色毒蛾核型多角体病毒(Dryobota furva nucleopolyhedrovirus) |
| 变形毒蛾核型多角体病毒(Dryobota protea nucleopolyhedrovirus) |
| 单色卓夜蛾核型多角体病毒(Dryobotodes monochroma nucleopolyhedrovirus) |
| 埃及金刚钻核型多角体病毒(Earias insulana nucleopolyhedrovirus) |
| 冰灯蛾核型多角体病毒(Ecpantheria icasia nucleopolyhedrovirus) |
| 鞍形尺蛾核型多角体病毒(Ectropis crepuscularia nucleopolyhedrovirus) |
| 茶尺蠖核型多角体病毒(Ectropis obliqua nucleopolyhedrovirus) |
| 橡尺蠖核型多角体病毒(Ennomos quercaria nucleopolyhedrovirus) |
| 栎尺蠖核型多角体病毒(Ennomos quercinaria nucleopolyhedrovirus) |
| 榆角尺蠖核型多角体病毒(Ennomos subsignarius nucleopolyhedrovirus) |
| 南加拿大尺蠖核型多角体病毒(Enypia venata nucleopolyhedrovirus) |
| 美洲银星弄蝶核型多角体病毒(Epargyreus clarus nucleopolyhedrovirus) |
| 烟草粉斑螟核型多角体病毒(Ephestia elutella nucleopolyhedrovirus) |
| 苹浅褐卷蛾核型多角体病毒(Epiphyas postvittana nucleopolyhedrovirus) |
| 落叶松尺蠖核型多角体病毒(Erannis ankeraria nucleopolyhedrovirus) |
| 暗点褚尺蠖核型多角体病毒(Erannis defoliaria nucleopolyhedrovirus) |
| 菩提尺蠖核型多角体病毒(Erannis tiliaria nucleopolyhedrovirus) |
| 多萼淫羊霍尺蠖核型多角体病毒(Erannis vancouverensis nucleopolyhedrovirus) |
| 木薯天蚕核型多角体病毒(Eratmapodites quinquevittatus nucleopolyhedrovirus) |
| 木薯天蛾核型多角体病毒(Erinnyis ello nucleopolyhedrovirus) |
| 樟蚕核型多角体病毒(Eriogyna pyretorum nucleopolyhedrovirus) |
| 盐泽枝灯蛾核型多角体病毒(Estigmene acrea nucleopolyhedrovirus) |
| 环形尺蠖核型多角体病毒(Eupithecia annulata nucleopolyhedrovirus) |
| 长足尺蠖核型多角体病毒(Eupithecia longipalpata nucleopolyhedrovirus) |
| 乌桕黄毒蛾核型多角体病毒(Euproctis bipunctapex nucleopolyhedrovirus) |
| 黄毒蛾核型多角体病毒(Euproctis chrysorrhoea nucleopolyhedrovirus) |
| 折带黄毒蛾核型多角体病毒(Euproctis flava nucleopolyhedrovirus) |
| 星黄毒蛾核型多角体病毒(Euproctis flavinata nucleopolyhedrovirus) |
| 缀黄毒蛾核型多角体病毒(Euproctis karghalica nucleopolyhedrovirus) |
| 茶黄毒蛾核型多角体病毒(Euproctis pseudoconspersa nucleopolyhedrovirus) |
| 盗毒蛾核型多角体病毒(Euproctis similis nucleopolyhedrovirus) |
| 折带黄毒蛾核型多角体病毒(Euproctis subflava nucleopolyhedrovirus) |
| 瑞木波纹蛾核型多角体病毒(Euthyatira pudens nucleopolyhedrovirus) |
| 增切根虫核型多角体病毒(Euxoa auxiliaris nucleopolyhedrovirus) |
| 暗缘地老虎核型多角体病毒(Euxoa messoria nucleopolyhedrovirus) |
| 红背地老虎核型多角体病毒(Euxoa ochrogaster nucleopolyhedrovirus) |
| 白地老虎核型多角体病毒(Euxoa scandens nucleopolyhedrovirus) |
| 野冬夜蛾核型多角体病毒(Feralia jacosa nucleopolyhedrovirus) |
| 李枯叶蛾核型多角体病毒(Gastropacha quercifolia nucleopolyhedrovirus) |
| 污行军虫核型多角体病毒(Hadena sordida nucleopolyhedrovirus) |
| 银点灯蛾核型多角体病毒(Halisidota argentata nucleopolyhedrovirus) |
| 胡桃点灯蛾核型多角体病毒(Halisidota caryae nucleopolyhedrovirus) |
| 菸夜蛾核型多角体病毒(Helicoverpa assulta nucleopolyhedrovirus) |
| 菜豆夜蛾核型多角体病毒(Helicoverpa obtectus nucleopolyhedrovirus) |
| 奇异夜蛾核型多角体病毒(Helicoverpa paradoxa nucleopolyhedrovirus) |
| 大棉铃虫核型多角体病毒(Helicoverpa peltigera nucleopolyhedrovirus) |
| 红噬斑夜蛾核型多角体病毒(Helicoverpa phloxiphaga nucleopolyhedrovirus) |
| 澳洲棉铃虫核型多角体病毒(Helicoverpa punctigera nucleopolyhedrovirus) |
| 红毒扁豆夜蛾核型多角体病毒(Helicoverpa rubrescens nucleopolyhedrovirus) |
| 亚曲夜蛾核型多角体病毒(Helicoverpa subflexa nucleopolyhedrovirus) |
| 烟芽夜蛾核型多角体病毒(Helicoverpa virescens nucleopolyhedrovirus) |
| 痣鳞褐蛉核型多角体病毒(Hemerobius stigma nucleopolyhedrovirus) |
| 赤杨条纹叶蜂核型多角体病毒(Hemichroa crocea nucleopolyhedrovirus) |
| 无背腺大蚕蛾核型多角体病毒(Hemileuca eglanterina nucleopolyhedrovirus) |
| 雄鹿大蚕蛾核型多角体病毒(Hemileuca maia nucleopolyhedrovirus) |
| 行列大蚕蛾核型多角体病毒(Hemileuca oliviae nucleopolyhedrovirus) |
| 三色大蚕蛾核型多角体病毒(Hemileuca tricolor nucleopolyhedrovirus) |
| 粉蝶蛾核型多角体病毒(Hesperumia sulphuraria nucleopolyhedrovirus) |
| 内斜线天蛾核型多角体病毒(Hippotion eson nucleopolyhedrovirus) |
| 茶长卷蛾核型多角体病毒(Homona magnanima nucleopolyhedrovirus) |
| 四叉莜夜蛾核型多角体病毒(Hoplodrina ambigua nucleopolyhedrovirus) |
| 惜古比天蚕核型多角体病毒 (Hyalophora cecropia nucleopolyhedrovirus) |
| Hyeriomena irata核型多角体病毒(Hyeriomena irata nucleopolyhedrovirus) |
| Hyeriomena nubilofasciata核型多角体病毒(Hyeriomena nubilofasciata nucleopolyhedrovirus) |
| 大戟天蛾核型多角体病毒(Hyles euphorbiae nucleopolyhedrovirus) |
| 五倍子天蛾核型多角体病毒(Hyles gallii nucleopolyhedrovirus) |
| 白线天蛾核型多角体病毒(Hyles lineata nucleopolyhedrovirus) |
| 黑棘天蛾核型多角体病毒(Hylesia nigricans nucleopolyhedrovirus) |
| 松尺蛾核型多角体病毒(Hyloicus pinastri nucleopolyhedrovirus) |
| 美洲螟核型多角体病毒(Hyperetis amicaria nucleopolyhedrovirus) |
| 石莲子天蛾核型多角体病毒(Hyphorma minax nucleopolyhedrovirus) |
| 火燕兰天蚕核型多角体病毒(Hypocrita jacobeae nucleopolyhedrovirus) |
| 孔雀翎蛱蝶核型多角体病毒(Inachis io nucleopolyhedrovirus) |
| 横带突刺蛾核型多角体病毒(Iragoides fasciata nucleopolyhedrovirus) |
| 黄足毒蛾核型多角体病毒(Ivela auripes nucleopolyhedrovirus) |
| 榆黄足毒蛾核型多角体病毒(Ivela ochropoda nucleopolyhedrovirus) |
| 运动尺蛾核型多角体病毒(Jankowskia athleta nucleopolyhedrovirus) |
| 聚蛱蝶核型多角体病毒(Junonia coenia nucleopolyhedrovirus) |
| 番茄夜蛾核型多角体病毒(Lacanobia oleracea nucleopolyhedrovirus) |
| 东方铁杉尺蠖核型多角体病毒(Lambdina fiscellaria nucleopolyhedrovirus) |
| 白杨天蛾核型多角体病毒(Laothoe populi nucleopolyhedrovirus) |
| 栎枯叶蛾核型多角体病毒(Lasiocampa quercus nucleopolyhedrovirus) |
| 三叶枯叶蛾核型多角体病毒(Lasiocampa trifolii nucleopolyhedrovirus) |
| 油茶大毛虫核型多角体病毒(Lebeda nobilis nucleopolyhedrovirus) |
| 枯叶蛾核型多角体病毒(Lechriolepis basirufa nucleopolyhedrovirus) |
| 杨雪毒蛾核型多角体病毒(Leucoma candida nucleopolyhedrovirus) |
| 雪毒蛾核型多角体病毒(Leucoma salicis nucleopolyhedrovirus) |
| 桦羽翅舟蛾核型多角体病毒(Lophopteryx camelina nucleopolyhedrovirus) |
| 甜菜网野螟核型多角体病毒(Loxostege sticticalis nucleopolyhedrovirus) |
| 日本虎凤蝶核型多角体病毒(Luehdorfia japonica nucleopolyhedrovirus) |
| 条毒蛾核型多角体病毒(Lymantria dissoluta nucleopolyhedrovirus) |
| 红腹毒蛾核型多角体病毒(Lymantria fumida nucleopolyhedrovirus) |
| 榄仁树毒蛾核型多角体病毒(Lymantria incerta nucleopolyhedrovirus) |
| 栎舞毒蛾核型多角体病毒(Lymantria mathura nucleopolyhedrovirus) |
| 模毒蛾核型多角体病毒(Lymantria monacha nucleopolyhedrovirus) |
| 九纹毒蛾核型多角体病毒(Lymantria ninayi nucleopolyhedrovirus) |
| 苹舞毒蛾核型多角体病毒(Lymantria obfuscata nucleopolyhedrovirus) |
| 红缘舞毒蛾核型多角体病毒(Lymantria violas winhol nucleopolyhedrovirus) |
| 木毒蛾核型多角体病毒(Lymantria xylina nucleopolyhedrovirus) |
| 杂灌枯叶蛾核型多角体病毒(Macrothylacia rubi nucleopolyhedrovirus) |
| 茶褐蓑蛾核型多角体病毒(Mahasena miniscula nucleopolyhedrovirus) |
| 阿尔俾天幕毛虫核型多角体病毒(Malacosoma alpicola nucleopolyhedrovirus) |
| 东方天幕毛虫核型多角体病毒(Malacosoma americanum nucleopolyhedrovirus) |
| 加州天幕毛虫核型多角体病毒(Malacosoma californicum nucleopolyhedrovirus) |
| 收缩天幕毛虫核型多角体病毒(Malacosoma constrictum nucleopolyhedrovirus) |
| 森林天幕毛虫核型多角体病毒(Malacosoma disstria nucleopolyhedrovirus) |
| 柔天幕毛虫核型多角体病毒(Malacosoma fragile nucleopolyhedrovirus) |
| 草原天幕毛虫核型多角体病毒(Malacosoma lutescens nucleopolyhedrovirus) |
| 黄褐天幕毛虫核型多角体病毒(Malacosoma neustria nucleopolyhedrovirus) |
| 西部天幕毛虫核型多角体病毒(Malacosoma pluviale nucleopolyhedrovirus) |
| 俗灰夜蛾核型多角体病毒(Mamestra suasa nucleopolyhedrovirus) |
| 烟草天蛾核型多角体病毒(Manduca sexta nucleopolyhedrovirus) |
| 绿纹森林活套蛾核型多角体病毒(Melanolophia imitata nucleopolyhedrovirus) |
| 狄网蛱蝶核型多角体病毒(Melitaea didyma nucleopolyhedrovirus) |
| 金钱松小卷蛾核型多角体病毒(Merophyas divulsana nucleopolyhedrovirus) |
| 樟叶蜂核型多角体病毒(Mesoneura rufonota nucleopolyhedrovirus) |
| 镶缤夜蛾核型多角体病毒(Moma champa nucleopolyhedrovirus) |
| 清波皎尺蛾核型多角体病毒(Myrteta tinagmaria nucleopolyhedrovirus) |
| 八纹噬叶野螟核型多角体病毒(Nacoleia octosema nucleopolyhedrovirus) |
| 白点舟蛾核型多角体病毒(Nadata gibbosa nucleopolyhedrovirus) |
| 美洲松粉蝶核型多角体病毒(Neophasia menapia nucleopolyhedrovirus) |
| 云舟蛾核型多角体病毒(Neopheosia excurvata nucleopolyhedrovirus) |
| 青铜地老虎核型多角体病毒(Nephelodes emmedonia nucleopolyhedrovirus) |
| 西方拟铁杉尺蠖核型多角体病毒(Nepytia freemani nucleopolyhedrovirus) |
| 铁杉尺蠖核型多角体病毒(Nepytia phantasmaria nucleopolyhedrovirus) |
| 夜蛾核型多角体病毒(Noctua pronuba nucleopolyhedrovirus) |
| 黄线森林尺蛾核型多角体病毒(Nyctobia limitaria nucleopolyhedrovirus) |
| 黄缘蛱蝶核型多角体病毒(Nymphalis antiopa nucleopolyhedrovirus) |
| 蛱蝶核型多角体病毒(Nymphalis polycholoros nucleopolyhedrovirus) |
| 稻三点水螟核型多角体病毒(Nymphula depunctalis nucleopolyhedrovirus) |
| 灰白蚕蛾核型多角体病毒(Ocinara varians nucleopolyhedrovirus) |
| 幽波尺蛾核型多角体病毒(Operophtera bruceata nucleopolyhedrovirus) |
| 冬尺蠖蛾核型多角体病毒(Operophtera brumata nucleopolyhedrovirus) |
| 热带蛾核型多角体病毒(Opisina arenosella nucleopolyhedrovirus) |
| 黄尺蛾核型多角体病毒(Opisthograpus luteolata nucleopolyhedrovirus) |
| 秋峦冬夜蛾核型多角体病毒(Oporinia autumnata nucleopolyhedrovirus) |
| 裂带蛾核型多角体病毒(Opsiphanes cassina nucleopolyhedrovirus) |
| 嘴壶夜蛾核型多角体病毒(Oraesia emarginata nucleopolyhedrovirus) |
| 澳古毒蛾核型多角体病毒(Orgyia anartoides nucleopolyhedrovirus) |
| 古毒蛾核型多角体病毒(Orgyia antiqua nucleopolyhedrovirus) |
| 澳洲古毒蛾核型多角体病毒(Orgyia australis nucleopolyhedrovirus) |
| 栗色古毒蛾核型多角体病毒(Orgyia badia nucleopolyhedrovirus) |
| 角斑古毒蛾核型多角体病毒(Orgyia gonostigma nucleopolyhedrovirus) |
| 合毒蛾核型多角体病毒(Orgyia leucostigma nucleopolyhedrovirus) |
| 棉古毒蛾核型多角体病毒(Orgyia postica nucleopolyhedrovirus) |
| 环古毒蛾核型多角体病毒(Orgyia turbata nucleopolyhedrovirus) |
| 老古毒蛾核型多角体病毒(Orgyia vetusta nucleopolyhedrovirus) |
| 冬夜蛾核型多角体病毒(Orthosia hibisci nucleopolyhedrovirus) |
| 云斑褐夜蛾核型多角体病毒(Orthosia incerta nucleopolyhedrovirus) |
| 玉米螟核型多角体病毒(Ostrinia nubilalis nucleopolyhedrovirus) |
| 佛得角枯叶蛾核型多角体病毒(Pachypasa capensis nucleopolyhedrovirus) |
| 奥特斯枯叶蛾核型多角体病毒(Pachypasa otus nucleopolyhedrovirus) |
| 春尺蠖核型多角体病毒(Paleacrita vernata nucleopolyhedrovirus) |
| 猩红虎蛾核型多角体病毒(Panaxia dominula nucleopolyhedrovirus) |
| 苹褐卷蛾核型多角体病毒(Pandemis heparana nucleopolyhedrovirus) |
| 栎褐卷蛾核型多角体病毒(Pandemis lamprosana nucleopolyhedrovirus) |
| 刚竹毒蛾核型多角体病毒(Pantana phyllostachysae nucleopolyhedrovirus) |
| 黑星剑纹夜蛾核型多角体病毒(Panthea portlandia nucleopolyhedrovirus) |
| 褐边绿刺蛾核型多角体病毒(Parasa consocia nucleopolyhedrovirus) |
| 丽绿刺蛾核型多角体病毒(Parasa lepida nucleopolyhedrovirus) |
| 中国绿刺蛾核型多角体病毒(Parasa sinica nucleopolyhedrovirus) |
| 直纹稻苞虫核型多角体病毒(Parnara guttata nucleopolyhedrovirus) |
| 隐纹谷弄蝶核型多角体病毒(Parnara mathias nucleopolyhedrovirus) |
| 线道尼斯凤蝶核型多角体病毒(Papilio daunis nucleopolyhedrovirus) |
| 达摩凤蝶核型多角体病毒(Papilio demoleus nucleopolyhedrovirus) |
| 杏凤蝶核型多角体病毒(Papilio podalirius nucleopolyhedrovirus) |
| 北美黑凤蝶核型多角体病毒(Papilio polyxenes nucleopolyhedrovirus) |
| 花椒凤蝶核型多角体病毒(Papilio xuthus nucleopolyhedrovirus) |
| 棉红铃虫核型多角体病毒(Pectinophora gossypiella nucleopolyhedrovirus) |
| 单尺蛾核型多角体病毒(Peribatoides simpliciaria nucleopolyhedrovirus) |
| 蓖麻斑灯蛾核型多角体病毒(Pericallia ricini nucleopolyhedrovirus) |
| 杂色地老虎核型多角体病毒(Peridroma saucia nucleopolyhedrovirus) |
| 巴西尺蠖核型多角体病毒( Pero behiensarius nucleopolyhedrovirus) |
| 米宙尺蠖核型多角体病毒(Pero mizon nucleopolyhedrovirus) |
| 栎掌舟蛾核型多角体病毒(Phalera assimilis nucleopolyhedrovirus) |
| 圆掌舟蛾核型多角体病毒(Phalera bucephala nucleopolyhedrovirus) |
| 大蚕蛾核型多角体病毒(Saturnia pyri nucleopolyhedrovirus) |
6 甲型杆状病毒属基本特性
核型多角体病毒的包涵体称多角体(polyhedron)。多角体的形状有三角形、四角形、五角形、六角形、立方形、近圆形和不规则形等。多角体的大小,一般变动在0.5~15µm之间,大多数为0.6~2.5µm。
多角体的表面具有多层结构的外膜,称多角体膜。膜内为多角体基质蛋白,由大分子的晶格组成,蛋白质分子呈立方形体系排列,基质蛋白称多角体蛋白(Polyhedrin)。多角体蛋白包埋着大量的病毒粒子或病毒束,病毒粒子或病毒束被包埋于多角体内是随机分布的,并不干扰多角体蛋白的晶格图像。多角体不溶于水、甲醛、乙醇、二甲苯、乙醚、1mol/L盐酸,0.75%胰蛋白酶,但0.1mol/L碳酸钠能使多角体溶解。
核型多角体病毒粒子是由衣壳(Capsid)包裹着髓核(Core)构成核衣壳(Nucleocapsid),核衣壳外被有囊膜(Envelope)所组成。病毒粒子呈杆状,其大小为40nm ~70nm×250 nm ~400nm,而核衣壳一般长度为300 nm ~330nm,直径35 nm~45nm。
核型多角体病毒有两个类型:单核衣壳多角体病毒(Single nucleocapsid polyhedrosis virus,SNPV)和多核衣壳多角体病毒(Multiple nucleocapsid polyhedrosis virus,MNPV)。单核衣壳多角体病毒是在一个囊膜内只含有一个核衣壳,如油桐尺蠖核型多角体病毒等;而多核衣壳多角体病毒的特点是在一个囊膜内含有2个以上的核衣壳,由多个核衣壳成束地被包装在同一个病毒囊膜内,又称病毒束(Virus bundle)。不同的毒株,MNPV的囊膜内包被的核衣壳数目是不同的,如:干果斑螟核型多角体病毒,病毒囊膜内包含的核衣壳数变动在2~23个,苜蓿银纹夜蛾核型多角体病毒,囊膜内包含核衣壳数变动于1~17个,而桑毛虫核型多角体病毒,一个囊膜包被的核衣壳数变动在1~39个。囊膜内的核衣壳大多数是有规律的排列的,如囊膜内含有3个核衣壳的,其排列图式呈三角形,囊内含有4个核衣壳的,其排列图式呈菱形,囊膜内含有5个核衣壳的,呈梯形排列图式或呈五角形排列。有时也观察到少数的囊膜内核衣壳呈不规则排列现象。
苜蓿银纹夜蛾NPV是第一个完成全序列的杆状病毒,现已有多种杆状病毒完成全序列分析。全世界各地从昆虫中分离的核型多角体病毒500多个,大部分从鳞翅目昆虫中分离而得,另外膜翅目31个,双翅目27个,鞘翅目5个,脉翅目2个,蚤目、缨尾目、毛翅目各1个。